Prof. Dr. Stefan Garthe

Hafentörn 1, D-25761 Büsum
Telefon: +49 4834 604-116
Telefax: +49 4834 604-299

Curriculum Vitae

Prof. Dr. Stefan Garthe


  • Seit 2014: Leitung der Arbeitsgruppe Ökologie mariner Tiere
  • 2014: Ernennung zum Professor an der Christian-Albrechts-Universität zu Kiel
  • 2001-2014: Leitung der Arbeitsgruppe Seevogelökologie
  • Seit 2004 außerdem Adjunct Professor am Department of Psychology an der Memorial University of Newfoundland, St. John's, Kanada
  • 1997-2001: Wissenschaftlicher Angestellter am Institut für Meereskunde an der Universität Kiel
  • 2005: Habilitation in Zoologie und Biologische Meereskunde an der Christian-Albrechts-Universität zu Kiel; Titel der Habilitationsschrift "Distribution, activity and foraging behaviour of seabirds in north-central European seas in relation to physical, biological and anthropogenic conditions"
  • 1997: Promotion in Zoologie an der Christian-Albrechts-Universität zu Kiel; Titel der Dissertation "Distribution and abundance of North Sea seabirds and their feeding ecology in relation to fisheries and hydrography"
  • 1993: Diplom in Biologie an der Christian-Albrechts-Universität zu Kiel (Hauptfach: Zoologie, Nebenfächer: Biologische Meereskunde und Physikalische Ozeanographie); Titel der Diplomarbeit "Quantifizierung von Abfall und Beifang der Fischerei in der südöstlichen Nordsee und deren Nutzung durch Seevögel"


  • Untersuchungen zur Ernährungsökologie ausgewählter Seevogelarten durch die Anwendung von Datenlogger-Technologie (Telemetrie) und Nahrungsanalysen
  • Räumlich-zeitliche Dynamik von Seevögeln in Schelfgebieten
  • Zusammenhänge zwischen abiotischen und biotischen Faktoren und Seevogel-Verteilungsmustern
  • Kurz- und langfristige Bestandsänderungen bei Seevögeln und ihre Ursachen
  • Wissenschaftliche Grundlagen im Konfliktfeld zwischen Meeresnaturschutz und Meeresnutzung (vor allem Fischerei und Offshore-Windkraft)


Master of Science Programm "Biological Oceanography":

  • "Seabird ecology" (Modul MNF-bioc-264; Praktikum)

Master of Science Programm "Environmental Management":

  • "Marine Nature Conservation: Theory and application" (Modul EMFTZ-01a; Vorlesung, Seminar, Übung)
  • "Abiotic-biotic interactions in the Wadden Sea" (Modul S 125; Vorlesung, Übung)
  • "Marine Ecology" als Teil von "Principles in Ecology for Environmental Sciences" (Modul S 154; Vorlesung, Übung)




Alle > 270 Publikationen sind hier vollständig gelistet.

Nachfolgend eine Auswahl:

  • Mercker M, Dierschke V, Camphuysen K, Kreutle A, Markones N, Vanermen N, Garthe S (im Druck): An indicator for assessing the status of marine-bird habitats affected by multiple human activities: A novel statistical approach. Ecological Indicators
  • Amélineau F, Delbart N, Schwemmer P, Marja R, Fort J, Garthe S, Elts J, Delaporte P, Rousseau P, Duraffour F, Bocher P (im Druck): Long distance migrants synchronize their spring departure with previous year conditions at their breeding site. Biology Letters
  • Enners L, Mercker M, Schwemmer P, Horn S, Asmus R, Asmus H, Eskildsen K, Schückel U, Garthe S (2021): Suitability of herring gulls (Larus argentatus) as indicators for detecting intertidal bivalve beds in the Wadden Sea. Ecological Indicators 129: 107947
  • Mercker M, Schwemmer P, Enners L, Peschko V, Garthe S (2021): Analysis of local habitat selection and large-scale attraction/avoidance based on animal tracking data: is there a single best method? Movement Ecology 9: 20.
  • Schwemmer P, Weiel S, Garthe S (2021): Spatio-temporal movement patterns and habitat choice of red foxes (Vulpes vulpes) and racoon dogs (Nyctereutes procyonoides) along the Wadden Sea coast. European Journal of Wildlife Research 67: 49.
  • Mercker M, Markones N, Borkenhagen K, Schwemmer H, Wahl J, Garthe S (2021): An integrated framework to estimate seabird population numbers and trends. Journal of Wildlife Management 1-21.
  • Schwemmer P, Mercker M, Vanselow KH, Bocher P, Garthe S (2021): Migrating curlews on schedule: departure and arrival patterns of a long-distance migrant depend on time and breeding location rather than on wind conditions. Movement Ecology 9: 9
  • Borrmann RM, Phillips RA, Clay TA, Garthe S (2021): Post-fledging migration and wintering strategies of individual juvenile Lesser Black-backed Gulls (Larus fuscus). Ibis 163: 1017-1031
  • Peschko V, Mendel B, Mercker M, Dierschke J, Garthe S (2021): Northern gannets (Morus bassanus) are strongly affected by operating offshore wind farms during the breeding season. Journal of Environmental Management 279: 111509
  • Lerma M, Dehnhard N, Luna-Jorquera G, Voigt CC, Garthe S (2020): Breeding stage, not sex, affects foraging characteristics in masked boobies at Rapa Nui. Behavioral Ecology and Sociobiology 74: 149.
  • Davidson SC, Bohrer G, … Garthe S, … (147 Autoren) (2020): Ecological insights from three decades of animal movement tracking across a changing Arctic. Science 370: 712–715
  • Peschko V, Mendel B, Müller S, Markones N, Mercker M, Garthe S (2020): Effects of offshore windfarms on seabird abundance: strong effects in spring and in the breeding season. Marine Environmental Research 162: 105157
  • Peschko V, Müller S, Schwemmer P, Mercker M, Lienau P, Rosenberger T, Sundermeyer J, Garthe S (2020): Wide dispersal of recently weaned grey seal pups in the southern North Sea. ICES Journal of Marine Science 77: 1762-1771
  • Enners L, Guse N, Schwemmer P, Chagas ALJ, Voigt CC, Garthe S (2020): Foraging ecology and diet of Eurasian spoonbills (Platalea leucorodia) in the German Wadden Sea. Estuarine, Coastal and Shelf Science 233: 106539.
  • Sherley RB, Ladd-Jones H, Garthe S, Stevenson O, Votier SC (2020): Scavenger communities and fisheries waste: North Sea discards support 3 million seabirds, 2 million fewer than in 1990. Fish and Fisheries 21: 132-145.
  • Schwemmer P, Adler S, Enners L, Volmer H, Kottsieper J, Ricklefs K, Stage M, Schwarzer K, Wittbrodt K, Reimers H-C, Binder K, Asmus R, Asmus H, Horn S, Schückel U, Kohlus J, Eskildsen K, Klingbeil K, Gräwe U, Garthe S (2019): Modelling and predicting habitats for the neobiotic American razor clam Ensis leei in the Wadden Sea. Estuarine, Coastal and Shelf Science 231: 106440.
  • Fliessbach KL, Borkenhagen K, Guse N, Markones N, Schwemmer P, Garthe S (2019): A ship traffic disturbance vulnerability index for Northwest European seabirds as a tool for Marine Spatial Planning. Frontiers in Marine Science 6: 192. doi: 10.3389/fmars.2019.00192
  • Mendel B, Schwemmer P, Peschko V, Müller S, Schwemmer H, Mercker M, Garthe S (2019): Operational offshore wind farms and associated ship traffic cause profound changes in distribution patterns of loons (Gavia spp.). Journal of Environmental Management 231: 429-438.
  • Corman A-M, Schwemmer P, Mercker M, Asmus H, Rüdel H, Klein R, Boner M, Hofem S, Koschorreck J, Garthe S (2018): Decreasing δ13C and δ15N values in four coastal species at different trophic levels indicate a fundamental food-web shift in the southern North and Baltic Seas between 1988 and 2016. Environmental Monitoring and Assessment 190: 461.
  • Enners L, Schwemmer P, Corman AM, Voigt CC, Garthe S (2018): Intercolony variations in movement patterns and foraging behaviors among herring gulls (Larus argentatus) breeding in the eastern Wadden Sea. Ecology and Evolution 8: 7529-7542.
  • Borkenhagen K, Corman A-M, Garthe S (2018): Estimating flight heights of seabirds using optical rangefinders and GPS data loggers: a methodological comparison. Marine Biology 165: 17.
  • Busch M, Garthe S (2018): Looking at the bigger picture: The importance of considering annual cycles in impact assessments illustrated in a migratory seabird species. ICES Journal of Marine Science 75: 690-700.
  • Garthe S, Peschko V, Kubetzki U, Corman A-M (2017): Seabirds as samplers of the marine environment – a case study in Northern Gannets. Ocean Science 13: 337-347.
  • Garthe S, Markones N, Corman A-M (2017): Possible impacts of offshore wind farms on seabirds: a pilot study in Northern Gannets in the southern North Sea. Journal of Ornithology 158: 345-349.
  • Dierschke V, Furness RW, Garthe S (2016): Seabirds and offshore wind farms in European waters: avoidance and attraction. Biological Conservation 202: 59-68.
  • Garthe S, Schwemmer P, Paiva VH, Corman A-M, Fock H, Voigt CC, Adler S (2016): Terrestrial and marine foraging strategies of an opportunistic seabird species breeding in the Wadden Sea. PLOS ONE 11(8): e0159630.
  • Garthe S, Hallgrimsson GT, Montevecchi WA, Fifield D, Furness RW (2016): East or west? Migration routes and wintering sites of Northern Gannets Morus bassanus from south-eastern Iceland. Marine Biology 163: 151.
  • Ramirez I, Paiva VH, Fagundes I, Menezes D, Silva I, Ceia FR, Phillips RA, Ramos JA, Garthe S (2016): Conservation implications of consistent foraging and trophic ecology in a rare petrel species. Animal Conservation 19: 139-152.
  • Sommerfeld J, Mendel B, Fock HO, Garthe S (2016): Combining bird-borne tracking and vessel-monitoring system data to assess discard use by a scavenging marine predator, the Lesser Black-backed Gull Larus fuscus. Marine Biology 163: 116.
  • Corman A-M, Mendel B, Voigt CC, Garthe S (2016): Varying foraging patterns in response to competition? A multicolony approach in a generalist seabird. Ecology and Evolution 6: 974-986.
  • Schwemmer P, Güpner F, Adler S, Klingbeil K, Garthe S (2016): Modelling small-scale foraging habitat use in Eurasian oystercatchers (Haematopus ostralegus) in relation to prey distribution and environmental predictors. Ecological Modelling 320: 322-333.
  • Schwemmer P, Covac A, Das K, Lepoint G, Adler S, Garthe S (2015): Assessment of contaminant levels and trophic relations at a World Heritage Site by measurements in a characteristic shorebird species. Environmental Research 136: 163-172.
  • Sommerfeld J, Kato A, Ropert-Coudert Y, Garthe S, Wilcox C, Hindell MA (2015): Flexible foraging behaviour in a marine predator according to foraging locations and environmental conditions. Journal of Experimental Marine Biology and Ecology 463: 79-86.
  • Corman A-M, Garthe S (2014): What flight heights tell us about foraging and potential conflicts with wind farms: a case study in Lesser Black-backed Gulls (Larus fuscus). Journal of Ornithology 155: 1037-1043.
  • Engelhard GH, Peck MA, Smout S, Rindorf A, Raab K, Aarts G, van Deurs M, Brunel T, Lauerburg RAM, Garthe S, Haste Andersen K, van Kooten T, Scott F, Dickey-Collas M (2014): Forage fish, their fisheries, and their predators: can we tell who drives whom in complex systems? ICES Journal of Marine Science 71: 90-104.
  • Garthe S,Guse N, Montevecchi WA,Rail JF, Gregoire F(2014): The daily catch: Flight altitude and diving behavior of northern gannets feeding on Atlantic mackerel. Journal of Sea Research 85: 456-462. doi:10.1016/j.seares.2013.07.020
  • Paiva VH, Geraldes P, Marques V, Rodríguez R, Ramírez I, Garthe S, Ramos JA (2013): Effects of environmental variability on different trophic levels of the North Atlantic food web. Marine Ecology Progress Series 477: 15-28.
  • Schwemmer H, Schwemmer P, Ehrich S, Garthe S (2013): Lesser black-backed gulls (Larus fuscus) consuming swimming crabs: An important link in the food web of the southern North Sea. Estuarine, Coastal and Shelf Science 119: 71-78.
  • Garthe S, Markones N, Mendel B, Sonntag N, Krause J (2012): Protected areas for seabirds in German offshore waters: designation, retrospective consideration and current perspectives. Biological Conservation 156: 126-135. doi:10.1016/j.biocon.2011.12.002
  • Garthe S, Ludynia K, Hüppop O, Kubetzki U, Meraz JF, Furness RW (2012): Energy budgets reveal equal benefits of varied migration strategies in northern gannets. Marine Biology 159: 1907–1915. doi:10.1007/s00227-012-1978-6
  • Fort J, Pettex E, Tremblay Y, Lorentsen S-H, Garthe S, Votier S, Pons JB, Siorat F, Furness RW, Grecian WJ, Bearhop S, Montevecchi WA, Grémillet D (2012): Meta-population evidence of oriented chain-migration in northern gannets. Frontiers in Ecology and the Environment 10: 237-242. doi:10.1890/110194
  • Montevecchi WA, Fifield D, Burke C, Garthe S, Hedd A, Rail J-F, Robertson G (2012): Tracking long-distance migration to assess marine pollution impact. Biology Letters 8: 218-221. doi:10.1098/rsbl.2011.0880
  • Sonntag N, Schwemmer H, Fock H, Bellebaum J, Garthe S (2012): Seabirds, set-nets, and conservation management: assessment of conflict potential and vulnerability of birds to bycatch in gillnets. ICES Journal of Marine Science 69: 578-589. doi:10.1093/icesjms/fss030
  • Garthe S, Montevecchi WA, Davoren GK (2011): Inter-annual changes in prey fields trigger different foraging tactics in a large marine predator. Limnology and Oceanography 56: 802-812. doi:10.4319/lo.2011.56.3.0802
  • Schwemmer P, Garthe S (2011): Spatial and temporal patterns of habitat use by Eurasian Oystercatchers (Haematopus ostralegus) population in the eastern Wadden Sea as revealed using GPS data loggers. Marine Biology 158: 541-550. doi:10.1007/s00227-010-1579-1
  • Schwemmer P, Mendel B, Sonntag N, Dierschke V, Garthe S (2011): Effects of ship traffic on seabirds in offshore waters: implications for marine conservation and spatial planning. Ecological Applications 21: 1851-1860. doi:10.1890/10-0615.1
  • Paiva VH, Geraldes P, Ramirez I, Garthe S, Ramos JA (2010): How area restricted search of a pelagic seabird changes while performing a dual foraging strategy. Oikos 119: 1423-1434. doi:10.1111/j.1600-0706.2010.18294.x
  • Garthe S, Markones N, Hüppop O, Adler S (2009): Effects of hydrographic and meteorological factors on seasonal seabird abundance in the southern North Sea. Marine Ecology Progress Series 391: 243-255. doi:10.3354/meps08170
  • Kubetzki U, Garthe S, Fifield D, Mendel B, Furness RW (2009): Individual migratory schedules and wintering areas of northern gannets. Marine Ecology Progress Series 391: 257-265. doi:10.3354/meps08254
  • Schwemmer P, Adler S, Guse N, Markones N, Garthe S (2009): Influence of water flow velocity and water depth on distribution and foraging patterns of a surface-feeding seabird in the German Wadden Sea. Fisheries Oceanography 18: 161-172. doi:10.1111/j.1365-2419.2009.00504.x
  • Zydelis R, Bellebaum J, Österblom H, Vetemaa M, Schirmeister B, Stipniece A, Dagys M, van Eerden M, Garthe S (2009): Bycatch in gillnet fisheries - An overlooked threat to waterbird populations. Biological Conservation 142: 1269-1281. doi:10.1016/j.biocon.2009.02.025
  • Schwemmer P, Garthe S, Mundry R (2008): Area utilization of gulls in a coastal farmland landscape: habitat mosaic supports niche segregation of opportunistic species. Landscape Ecology 23: 355-367. doi:10.1007/s10980-008-9194-y
  • Garthe S, Montevecchi WA, Chapdelaine G, Rail J-F, Hedd A (2007): Contrasting foraging tactics by northern gannets (Sula bassana) breeding in different oceanographic domains with different prey fields. Marine Biology 151: 687-694. doi:10.1007/s00227-006-0523-x
  • Garthe S, Montevecchi WA, Davoren GK (2007): Flight destinations and foraging behaviour of northern gannets (Sula bassana) preying on a small foraging fish in a low-Arctic ecosystem. Deep-Sea Research Part II 54: 311-320. doi:10.1016/j.dsr2.2006.11.008
  • Barrett RT, Camphuysen CJ, Anker-Nilssen T, Chardine JW, Furness RW, Garthe S, Hüppop O, Leopold MF, Montevecchi WA, Veit RR (2007): Diet studies of seabirds: a review and recommendations. ICES Journal of Marine Science 64: 1675-1691. doi:10.1093/icesjms/fsm152
  • Garthe S, Hüppop O (2004): Scaling possible adverse effects of marine wind farms on seabirds: developing and applying a vulnerability index. Journal of Applied Ecology 41: 724-734. doi:10.1111/j.0021-8901.2004.00918.x
  • Garthe S, Benvenuti S, Montevecchi WA (2003): Temporal patterns of foraging activities of northern gannets, Morus bassanus, in the northwest Atlantic Ocean. Canadian Journal of Zoology 81: 453-461. doi:10.1139/Z03-027
  • Garthe S, Scherp B (2003): Utilization of discards and offal from commercial fisheries by seabirds in the Baltic Sea. ICES Journal of Marine Science 60: 980-989. doi:10.1016/S1054-3139(03)00099-7
  • Kubetzki U, Garthe S (2003): Distribution, diet and habitat selection by four sympatrically breeding gull species in the southeastern North Sea. Marine Biology 143: 199-207. doi:10.1007/s00227-003-1036-5
  • Garthe S, Benvenuti S, Montevecchi WA (2000): Pursuit plunging by Northern Gannets (Sula bassana) feeding on Capelin (Mallotus villosus). Proceedings of the Royal Society London, Biological Sciences 267: 1717-1722. doi:10.1098/rspb.2000.1200
  • Tasker ML, Camphuysen CJ, Cooper J, Garthe S, Montevecchi WA, Blaber SJM (2000): The impacts of fishing on marine birds. ICES Journal of Marine Science 57: 531-547. doi:10.1006/jmsc.2000.0714
  • Garthe S, Grémillet D, Furness RW (1999): At-sea-activity and foraging efficiency in chick-rearing Northern Gannets (Sula bassana): a case study in Shetland. Marine Ecology Progress Series 185: 93-99. doi:10.3354/meps185093
  • Garthe S (1997): Influence of hydrography, fishing activity and colony location on summer seabird distribution in the southeastern North Sea. ICES Journal of Marine Science 54: 566-577. doi:10.1006/jmsc.1997.0253
  • Garthe S, Camphuysen CJ, Furness RW (1996): Amounts of discards by commercial fisheries and their significance as food for seabirds in the North Sea. Marine Ecology Progress Series 136: 1-11. doi:10.3354/meps136001
  • Garthe S, Hüppop O (1994): Distribution of ship-following seabirds and their utilization of discards in the North Sea in summer. Marine Ecology Progress Series 106: 1-9. (Paper)