ECOLAB - Group Marine Animal Ecology

In press

  • Amélineau F, Delbart N, Schwemmer P, Marja R, Fort J, Garthe S, Elts J, Delaporte P, Rousseau P, Duraffour F, Bocher P: Long distance migrants synchronize their spring departure with previous year conditions at their breeding site. Biology Letters
  • Mercker M, Dierschke V, Camphuysen K, Kreutle A, Markones N, Vanermen N, Garthe S: An indicator for assessing the status of marine-bird habitats affected by multiple human activities: A novel statistical approach. Ecological Indicators



  • Borrmann RM, Phillips RA, Clay TA, Garthe S (2021): Post-fledging migration and wintering strategies of individual juvenile Lesser Black-backed Gulls (Larus fuscus). Ibis 163: 1017-1031
  • Enners L, Mercker M, Schwemmer P, Horn S, Asmus R, Asmus H, Eskildsen K, Schückel U, Garthe S (2021): Suitability of herring gulls (Larus argentatus) as indicators for detecting intertidal bivalve beds in the Wadden Sea. Ecological Indicators 129: 107947
  • Mercker M, Markones N, Borkenhagen K, Schwemmer H, Wahl J, Garthe S (2021): An integrated framework to estimate seabird population numbers and trends. Journal of Wildlife Management 1-21.
  • Mercker M, Schwemmer P, Enners L, Peschko V, Garthe S (2021): Analysis of local habitat selection and large-scale attraction/avoidance based on animal tracking data: is there a single best method? Movement Ecology 9: 20.
  • Montevecchi WA, Regular PM, Rail J-F, Power K, Mooney C, D'Entremont KJN, Garthe S, Guzzwell L, Wilhelm S (2021): Ocean heat wave induces breeding failure at the southern breeding limit of the northern gannet Morus bassanus. Marine Ornithology 49: 71-78.
  • Peschko V, Mendel B, Mercker M, Dierschke J, Garthe S (2021): Northern gannets (Morus bassanus) are strongly affected by operating offshore wind farms during the breeding season. Journal of Environmental Management 279: 111509
  • Schwemmer P, Mercker M, Vanselow KH, Bocher P, Garthe S (2021): Migrating curlews on schedule: departure and arrival patterns of a long-distance migrant depend on time and breeding location rather than on wind conditions. Movement Ecology 9: 9
  • Schwemmer P, Weiel S, Garthe S (2021): Spatio-temporal movement patterns and habitat choice of red foxes (Vulpes vulpes) and racoon dogs (Nyctereutes procyonoides) along the Wadden Sea coast. European Journal of Wildlife Research 67: 49.



  • Davidson SC, Bohrer G, … Garthe S, …, Schwemmer P, … (147 authors) (2020): Ecological insights from three decades of animal movement tracking across a changing Arctic. Science 370: 712–715
  • Enners L, Guse N, Schwemmer P, Chagas ALJ, Voigt CC, Garthe S (2020): Foraging ecology and diet of Eurasian spoonbills (Platalea leucorodia) in the German Wadden Sea. Estuarine, Coastal and Shelf Science 233: 106539.
  • Horn S, Schwemmer P, Mercker M, Enners L, Asmus R, Garthe S, Asmus H (2020): Species composition of foraging birds in association with benthic fauna in four intertidal habitats of the Wadden Sea. Estuarine, Coastal and Shelf Science 233: 106537
  • Kubetzki U, Garthe S, Schwemmer P, Heinze B (2020): Sturmmöwe an der Ostsee: Küsten- oder Agrarvogel? Falke 67 (8): 8-13.
  • Lerma M, Castillo-Guerrero JA, Hernández-Vázquez S, Garthe S (2020): Foraging ecology of a marine top predator in the Eastern Tropical Pacific over 3 years with different ENSO phases. Marine Biology 167: 88.
  • Lerma M, Dehnhard N, Luna-Jorquera G, Voigt CC, Garthe S: Breeding stage, not sex, affects foraging characteristics in masked boobies at Rapa Nui. Behavioral Ecology and Sociobiology 74: 149
  • Lerma M, Serratosa J, Luna-Jorquera G, Garthe S (2020): Foraging ecology of masked boobies (Sula dactylatra) in the world’s largest "oceanic desert". Marine Biology 167: 87
  • Pelletier D, Seyer Y, Garthe S, Bonnefoi S, Phillips RA, Guillemette M (2020): So far, so good … Similar fitness consequences and overall energetic costs for short and long-distance migrants in a seabird. PLoS ONE 15(3): e0230262
  • Peschko V, Mendel B, Müller S, Markones N, Mercker M, Garthe S (2020): Effects of offshore windfarms on seabird abundance: strong effects in spring and in the breeding season. Marine Environmental Research 162: 105157
  • Peschko V, Mercker M, Garthe S (2020): Telemetry reveals strong effects of offshore wind farms on behaviour and habitat use of common guillemots (Uria aalge) during the breeding season. Marine Biology 167: 118.
  • Peschko V, Müller S, Schwemmer P, Mercker M, Lienau P, Rosenberger T, Sundermeyer J, Garthe S (2020): Wide dispersal of recently weaned grey seal pups in the southern North Sea. ICES Journal of Marine Science 77: 1762-1771
  • Sherley RB, Ladd-Jones H, Garthe S, Stevenson O, Votier SC (2020): Scavenger communities and fisheries waste: North Sea discards support 3 million seabirds, 2 million fewer than in 1990. Fish and Fisheries 21: 132-145.
  • Waggitt J, … Garthe S, … Markones N, … (41 authors) (2020): Distribution maps of cetacean and seabird populations in the North-East Atlantic. Journal of Applied Ecology 57: 253-269.



  • Borkenhagen K (2019): Erlenzeisige mögen Seafood - Nutzung einer ungewöhnlichen Nahrungsquelle bei ziehenden Singvögeln. Natur- und Landeskunde 126: 185-187.
  • Borkenhagen K, Markones N, Garthe S (2019): Das Wirbeltiermonitoring in den deutschen Meeresgebieten und die Rolle digitaler Erfassungen im Monitoringkonzept. Jahrbuch für Naturschutz und Landschaftspflege 62: 56-60.
  • Borrmann RM, Phillips RA, Clay T, Garthe S (2019): High foraging site fidelity and spatial segregation among individual great black-backed gulls. Journal of Avian Biology: e02156.
  • Enners L, Chagas ALJ, Schwemmer P, Ismar-Rebitz SMH, Garthe S (2019): Foraging patterns and diet composition of breeding pied avocets (Recurvirostra avosetta) in the German Wadden Sea. Estuarine, Coastal and Shelf Science 229: 106399.
  • Fliessbach KL, Borkenhagen K, Guse N, Markones N, Schwemmer P, Garthe S (2019): A ship traffic disturbance vulnerability index for Northwest European seabirds as a tool for Marine Spatial Planning. Frontiers in Marine Science 6: 192.
  • Horn S, de la Vega C, Asmus R, Schwemmer P, Enners L, Garthe S, Haslob H, Binder K, Asmus H (2019): Impact of birds on intertidal food webs assessed with ecological network analysis. Estuarine Coastal and Shelf Science 219: 107-119.
  • Kottsieper J, Schückel U, Schwemmer P, Fox AD, Garthe S (2019): Comparison of bivalve communities between moulting and wintering areas used by Common Scoter Melanitta nigra in the German North Sea. Estuarine, Coastal and Shelf Science 229: 106398.
  • Kottsieper J, Schwemmer P, Markones N, Fox AD, Garthe S (2019): An invasive alien bivalve apparently provides a novel food source for moulting and wintering benthic feeding sea ducks. Helgoland Marine Research 73: 11
  • Mendel B, Schwemmer P, Peschko V, Müller S, Schwemmer H, Mercker M, Garthe S (2019): Operational offshore wind farms and associated ship traffic cause profound changes in distribution patterns of loons (Gavia spp.). Journal of Environmental Management 231: 429-438.
  • Schwemmer P, Adler S, Enners L, Volmer H, Kottsieper J, Ricklefs K, Stage M, Schwarzer K, Wittbrodt K, Reimers H-C, Binder K, Asmus R, Asmus H, Horn S, Schückel U, Kohlus J, Eskildsen K, Klingbeil K, Gräwe U, Garthe S (2019): Modelling and predicting habitats for the neobiotic American razor clam Ensis leei in the Wadden Sea. Estuarine, Coastal and Shelf Science 231: 106440.
  • Schwemmer P, Corman A-M, Koch D, Borrmann RM, Koschorreck J, Voigt CC, Garthe S (2019): Charakterisierung der Habitatwahl von Silbermöwen Larus argentatus durch GPS-Datenlogger zur Einschätzung der Schadstoffbelastung an der deutschen Nordseeküste. Vogelwarte 57: 13-30.
  • Schwemmer P, Volmer H, Enners L, Reimers H-C, Binder K, Horn S, Adler S, Fox AD, Garthe S (2019): Modelling distribution of common scoter (Melanitta nigra) by its predominant prey, the American razor clam (Ensis leei) and hydrodynamic parameters. Estuarine, Coastal and Shelf Science 225: 106260



  • Borkenhagen K, Corman A-M, Garthe S (2018): Estimating flight heights of seabirds using optical rangefinders and GPS data loggers: a methodological comparison. Marine Biology 165: 17.
  • Busch M, Garthe S (2018): Looking at the bigger picture: The importance of considering annual cycles in impact assessments illustrated in a migratory seabird species. ICES Journal of Marine Science 75: 690-700.
  • Corman A-M, Schwemmer P, Mercker M, Asmus H, Rüdel H, Klein R, Boner M, Hofem S, Koschorreck J, Garthe S: Decreasing δ13C and δ15N values in four coastal species at different trophic levels indicate a fundamental food-web shift in the southern North and Baltic Seas between 1988 and 2016. Environmental Monitoring and Assessment 190: 461.
  • de la Vega C, Horn S, Baird D, Hines DE, Borrett S, Jensen L, Schwemmer P, Asmus R, Siebert U, Asmus H (2018) Seasonal dynamics and functioning of the Sylt-Rømø Bight, northern Wadden Sea. Estuarine Coastal and Shelf Science 203: 100-118.
  • Enners L, Schwemmer P, Corman AM, Voigt CC, Garthe S (2018): Intercolony variations in movement patterns and foraging behaviors among Herring Gulls (Larus argentatus) breeding in the eastern Wadden Sea. Ecology and Evolution 8: 7529-7542.
  • Furness RW, Garthe S, Trinder M, Matthiopoulos J, Wanless S, Jeglinski J (2018): Nocturnal flight activity of Northern Gannets Morus bassanus and implications for modelling collision risk at offshore wind farms. Environmental Impact Assessment Review 73: 1-6.
  • Furness RW, Hallgrimsson GT, Montevecchi WA, Fifield DA, Kubetzki U, Mendel B, Garthe S (2018): Adult Gannet migrations frequently loop clockwise around Britain and Ireland. Ringing and Migration 33: 45-53.
  • Guse N, Witte K, Markones N, Borkenhagen K, Scheiffarth G, Garthe S (2018): Aktuelle Verbreitung, Bestände und Trends von Seevögeln auf See im Offshore-Bereich des niedersächsischen Küstenmeers und des Nationalparks Niedersächsisches Wattenmeer. Vogelkdl. Ber. Niedersachsen 46: 1-69.



  • Baschek B, Schroeder F, Brix H, Riethmüller R, Badewien TH, Breitbach G, Brügge B, Colijn F, Doerffer R, Eschenbach C, Friedrich J, Fischer P, Garthe S, Horstmann J, Krasemann H, Metfies K, Ohle N, Petersen W, Pröfrock D, Röttgers R, Schlüter M, Schulz J, Schulz-Stellenfleth J, Stanev E, Winter C, Wirtz K, Wollschläger J, Zielinski O, Ziemer F (2017): The Coastal Observing System for Northern and Arctic Seas (COSYNA). Ocean Science 13: 379-410.
  • Borkenhagen K (2017) Molecular phylogeny of the tribe Torini Karaman, 1971 (Actinopterygii: Cypriniformes) from the Middle East and North Africa. Zootaxa 4236 (2): 291-301.
  • Busch M, Garthe S (2017): Looking at the bigger picture: The importance of considering annual cycles in impact assessments illustrated in a migratory seabird species. ICES Journal of Marine Science, doi:10.1093/icesjms/fsx170
  • Garthe S, Markones N, Corman A-M (2017): Possible impacts of offshore wind farms on seabirds: a pilot study in Northern Gannets in the southern North Sea. Journal of Ornithology 158: 345-349.
  • Garthe S, Peschko V, Kubetzki U, Corman A-M (2017): Seabirds as samplers of the marine environment – a case study in Northern Gannets. Ocean Science 13: 337-347.
  • Horn S, de la Vega C, Asmus R, Schwemmer P, Enners L, Garthe S, Binder K, Asmus H (2017): Interaction between birds and macrofauna within food webs of six intertidal habitats of the Wadden Sea. PLOS ONE 12(5): e0176381.
  • Ramírez I, Tarzia M, Dias M, Burfield I, Ramos J, Garthe S, Paiva VH (2017): How well is the EU protecting its seabirds? Progress in implementing the Birds Directive at sea. Marine Policy 81: 179-184.
  • Schwemmer P (2017) GPS-tracking of Eurasian Curlews and other coastal bird species indicate high connectivity between the German Wadden Sea and Russia (results of the German project „Birdmove“). Information Materials of the Working Group on Waders of Northern Eurasia 30: 46-48 (in Russian).
  • Schwemmer P, Schwemmer H, Garthe S (2017): Verbreitung, Phänologie und Habitatwahl von Möwen im Binnenland der Nordseeküste Schleswig-Holsteins, ermittelt durch Flugzeugerfassungen. Vogelwelt 137: 271-287.
  • Schwemmer P, Weiel S, Garthe S (2017): A fundamental study revisited: Quantitative evidence for territory quality in oystercatchers (Haematopus ostralegus) using GPS data loggers. Ecology and Evolution 7: 285-294.
  • Vanselow KH, Jacobsen S, Hall C, Garthe S (2017): Solar storms may trigger sperm whale strandings: explanation approaches for multiple strandings in the North Sea in 2016. International Journal of Astrobiology, doi:10.1017/S147355041700026X.



  • Brander KM, Ottersen G, Bakker JP, Beaugrand G, Herr H, Garthe S, Gilles A, Kenny A, Siebert U, Skjoldal HR, Tulp I (2016): Environmental Impacts - Marine Ecosystems. In: Quante M, Colijn F (eds): North Sea region climate change assessment. Regional Climate Studies. Springer. S. 241-274.
  • Busch M, Garthe S (2016): Approaching population thresholds in presence of uncertainty: Assessing displacement of seabirds from offshore wind farms. Environmental Impact Assessment Review 56: 31-24.
  • Castillo-Guerrero JA, Lerma M, Mellink E, Suazo-Guillen E, Peñaloza-Padilla EA (2016) Environmental-mediated flexible foraging strategies in Brown Boobies in the Gulf of California. Ardea 104: 33-47.
  • Corman A-M, Mendel B, Voigt CC, Garthe S (2016): Varying foraging patterns in response to competition? A multicolony approach in a generalist seabird. Ecology and Evolution 6: 974-986.
  • Dierschke V, Furness RW, Garthe S (2016): Seabirds and offshore wind farms in European waters: avoidance and attraction. Biological Conservation 202: 59-68.
  • Enners L, Schwemmer P, Garthe S (2016): Wo gibt's hier was zu essen? Ernährungsstrategien von Silbermöwen Larus argentatus, Lachmöwen L. ridibundus und Säbelschnäblern Recurvirostra avosetta an der schleswig-holsteinischen Nordseeküste. Vogelkundliche Berichte aus Niedersachsen 44: 323-324.
  • Garthe S, Hallgrimsson GT, Montevecchi WA, Fifield D, Furness RW (2016): East or west? Migration routes and wintering sites of Northern Gannets Morus bassanus from south-eastern Iceland. Marine Biology 163: 151.
  • Garthe S, Schwemmer P, Paiva VH, Corman A-M, Fock H, Voigt CC, Adler S (2016): Terrestrial and marine foraging strategies of an opportunistic seabird species breeding in the Wadden Sea. PLOS ONE 11(8): e0159630.
  • Ramirez I, Paiva VH, Fagundes I, Menezes D, Silva I, Ceia FR, Phillips RA, Ramos JA, Garthe S (2016): Conservation implications of consistent foraging and trophic ecology in a rare petrel species. Animal Conservation 19: 139-152.
  • Schwemmer P, Enners L, Garthe S (2016): Migration routes of Eurasian Curlews (Numenius arquata) resting in the eastern Wadden Sea based on GPS telemetry. Journal of Ornithology 157: 901-905.
  • Schwemmer P, Güpner F, Adler S, Klingbeil K, Garthe S (2016): Modelling small-scale foraging habitat use in Eurasian oystercatchers (Haematopus ostralegus) in relation to prey distribution and environmental predictors. Ecological Modelling 320: 322-333.
  • Schwemmer P, Voigt C, Corman A-M, Adler S, Garthe S (2016): Body mass change and diet switch tracked by stable isotopes indicate time spent at a stopover site during autumn migration in dunlins (Calidris alpina alpina). Journal of Avian Biology 47: 806-814.
  • Sommer F, Schwemmer P, Garthe S, Valqui J, Eckern S, Hartl GB (2016): Genetische Diversität des Sterntauchers Gavia stellata in der deutschen Nord- und Ostsee: Analyse der Domäne II der mitochondrialen Kontrollregion. Vogelkundliche Berichte aus Niedersachsen 44: 313-320.
  • Sommerfeld J, Mendel B, Fock HO, Garthe S (2016): Combining bird-borne tracking and vessel-monitoring system data to assess discard use by a scavenging marine predator, the Lesser Black-backed Gull Larus fuscus. Marine Biology 163: 116.
  • Volmer H, Schwemmer P, Garthe S (2016): Alle meine Entchen schwimmen auf der See – Aktuelles zur Nahrungswahl von Eider- Somateria mollissima und Trauerenten Melanitta nigra im schleswig-holsteinischen Wattenmeer. Vogelkundliche Berichte aus Niedersachsen 44: 321-322.



  • Ceia F, Paiva V, Ceia R, Hervías S, Garthe S, Marques J, Ramos J (2015): Spatial foraging segregation by close neighbours in a wide-ranging seabird. Oecologia 177: 431-440.
  • Garthe S, Schwemmer H, Markones N, Müller S, Schwemmer P (2015): Verbreitung, Jahresdynamik und Bestandsentwicklung der Seetaucher Gavia spec. in der Deutschen Bucht (Nordsee). Vogelwarte 53: 121-138.
  • Schwemmer P, Covac A, Das K, Lepoint G, Adler S, Garthe S (2015): Assessment of contaminant levels and trophic relations at a World Heritage Site by measurements in a characteristic shorebird species. Environmental Research 136: 163-172.
  • Sommerfeld J, Kato A, Ropert-Coudert Y, Garthe S, Wilcox C, Hindell MA (2015): Flexible foraging behaviour in a marine predator according to foraging locations and environmental conditions. Journal of Experimental Marine Biology and Ecology 463: 79-86.



  • Ceia F, Paiva V, Garthe S, Marques J, Ramos J (2014): Can variations in the spatial distribution at sea and isotopic niche width be associated with consistency in the isotopic niche of a pelagic seabird species? Marine Biology 161: 1861-1872.
  • Ceia FR, Paiva VH, Fidalgo V, Morais L, Baeta A, Crisóstomo P, Mourato E, Garthe S, Marques JC, Ramos JA (2014): Annual and seasonal consistency in the feeding ecology of an opportunistic species, the yellow-legged gull Larus michahellis. Marine Ecology Progress Series 497: 273-284
  • Corman A-M, Garthe S (2014): What flight heights tell us about foraging and potential conflicts with wind farms: a case study in Lesser Black-backed Gulls (Larus fuscus). Journal of Ornithology 155: 1037-1043.
  • Dickey-Collas M, Engelhard G, Rindorf A, Raab K, Smout S, Aarts G, van Deurs M, Brunel T, Hoff A, Lauerburg R, Garthe S, Haste Andersen K, Scott F, van Kooten T, Beare D, Peck M (2014): Ecosystem-based management objectives for the North Sea: Riding the Forage Fish Rollercoaster. ICES Journal of Marine Science 71: 128-142.
  • Engelhard GH, Peck MA, Smout S, Rindorf A, Raab K, Aarts G, van Deurs M, Brunel T, Lauerburg RAM, Garthe S, Haste Andersen K, van Kooten T, Scott F, Dickey-Collas M (2014): Forage fish, their fisheries, and their predators: can we tell who drives whom in complex systems? ICES Journal of Marine Science 71: 90-104.
  • Fifield DA, Montevecchi WA, Garthe S, Robertson GJ, Kubetzki U, Rail J-F (2014): Migratory tactics and wintering areas of Northern Gannets (Morus bassanus) breeding in North America. Ornithological Monographs 79: 1-63.
  • Garthe S, Guse N, Montevecchi WA, Rail J-F, Grégoire F (2014): The daily catch: Flight altitudes and diving behaviour of northern gannets feeding on Atlantic mackerel. Journal of Sea Research 85: 456-462.
  • Kubetzki U, Garthe S (2014): Neufundlands spektakuläre Vogelinsel Funk Island: Auf den Spuren des Riesenalks. Falke 61: 12-19.
  • Mendel B, Kotzerka J, Sommerfeld J, Schwemmer H, Sonntag N, Garthe S (2014): Effects of the alpha ventus offshore test site on distribution patterns, behaviour and flight heights of seabirds. In: Federal Maritime and Hydrographic Agency and Federal Ministry for the Environment, Nature Conservation and Nuclear Safety (eds.): Ecological Research at the Offshore Windfarm alpha ventus. Springer Spektrum, Wiesbaden. S. 95-110.
  • Schwemmer P, Hälterlein B, Geiter O, Günther K, Corman VM, Garthe S (2014): Weather-related winter mortality of Eurasian Oystercatchers (Haematopus ostralegus) in the northeastern Wadden Sea. Waterbirds 37: 319-330.
  • Wittwer C, Schwemmer P (2014): Einfluss von Eiswintern auf die Nahrungsgrundlage von Küstenvögeln im Dithmarscher Wattenmeer. Faunistisch‐ökologische Mitteilungen 9: 335‐356.



  • Bellebaum J, Schirmeister B, Sonntag N, Garthe S (2013) Decreasing but still high: Bycatch of seabirds in gillnet fisheries along the German Baltic coast. Aquatic Conservation: Marine and Freshwater Ecosystems 23: 210-221.
  • Busch M, Kannen A, Garthe S, Jessopp M (2013): Consequences of a cumulative perspective on marine environmental impacts: offshore wind farming and seabirds at North Sea scale in context of the EU Marine Strategy. Ocean & Coastal Management 71: 213-224.
  • Corman A-M, Garthe S (2013): Wo suchen die Heringsmöwen der Amrumer Odde ihre Nahrung? Seevögel 34: 156-159.
  • Guse N, Markones N, Bolduc F, Garthe S (2013): Distribution of seabirds in the Lower Estuary and Gulf of St Lawrence (Canada) during summer. Seabird 26: 42-70.
  • Kubetzki U (2013): Eine Erfolgsgeschichte: Datenlogger in der Seevogelforschung. Falke 60: 92-98. pdf zum Download (2 MB)
  • Kubetzki U, Schwemmer P (2013): Bestandsrückgänge des Austernfischers im Wattenmeer: Welche Rolle spielen Veränderungen im Nahrungshabitat? Falke 60: 371-375.
  • Montevecchi B, Chardine J, Rail J-F, Garthe S, Pelletier D, Regular P, Burke C, Hedd A, McFarlane Tranquilla L, Bennett S, Mooney C, Power K, Power T, Hogan H, Daoust P-Y, Lawson J, Rogers L, Wilhelm S, Montevecchi M, Lang A (2013): Extreme event in a changing ocean climate: warm-water perturbation of 2012 influences breeding gannets and other marine animals in the Northwest Atlantic and Gulf of St Lawrence. Osprey 44: 14-19.
  • Paiva VH, Geraldes P, Marques V, Rodríguez R, Ramírez I, Garthe S, Ramos JA (2013): Effects of environmental variability on different trophic levels of the North Atlantic food web. Marine Ecology Progress Series 477: 15-28.
  • Paiva VH, Geraldes P, Ramírez I, Werner AC, Garthe S, Ramos JA (2013): Overcoming difficult times: The behavioural resilience of a marine predator when facing environmental stochasticity. Marine Ecology Progress Series 486: 277-288.
  • Ramírez I, Paiva VH, Menezes D, Silva I, Phillips RA, Ramos JA, Garthe S (2013): Year-round distribution and habitat preferences of Bugio's petrel. Marine Ecology Progress Series 476: 269-284.
  • Schwemmer H, Schwemmer P, Ehrich S, Garthe S (2013): Lesser black-backed gulls (Larus fuscus) consuming swimming crabs: An important link in the food web of the southern North Sea. Estuarine, Coastal and Shelf Science 119: 71-78.
  • Sommerfeld J, Kato A, Ropert-Coudert Y, Garthe S, Hindell MA (2013): The individual counts: within sex differences in foraging strategies are as important as sex-specific differences in Masked Boobies Sula dactylatra. Journal of Avian Biology 44: 531-540.
  • Sommerfeld J, Kato A, Ropert-Coudert Y, Garthe S, Hindell MA (2013): Foraging parameters influencing the detection and interpretation of Area-restricted search behaviour in marine predators: a case study with the Masked Booby. PLoS One e63742.
  • Volmer H, Bickel M, Schwemmer P, Garthe S (2013): Kükenprädation bei Heringsmöwen auf Amrum als mögliche Erklärung für den schlechten Bruterfolg im Jahr 2012. Seevögel 34: 160-165.



  • Bond AL, Montevecchi WA, Guse N, Regular PM, Garthe S, Rail J-F (2012): Prevalence and composition of fishing gear debris in the nests of Northern Gannets (Morus bassanus) are related to fishing effort. Marine Pollution Bulletin 64: 907-911.
  • Camphuysen CJ, Shamoun-Baranes J, Bouten W, Garthe S (2012): Identifying ecologically important marine areas for seabirds using behavioural information in combination with distribution patterns. Biological Conservation 156: 22-29.
  • Dierschke V, Exo K-M, Mendel B, Garthe S (2012): Gefährdung von Sterntaucher Gavia stellata und Prachttaucher G. arctica in Brut-, Zug- und Überwinterungsgebieten – eine Übersicht mit Schwerpunkt auf den deutschen Meeresgebieten. Vogelwelt 133: 163-194.
  • Ekroos J, Fox AD, Christensen TK, Petersen IK, Kilpi M, Jónsson JE, Green M, Laursen K, Cervencl A, de Boer P, Nilsson L, Meissner W, Garthe S, Öst M (2012): Declines amongst breeding Eider Somateria mollissima numbers in the Baltic/Wadden Sea flyway. Ornis Fennica 89: 81-90.
  • Fort J, Pettex E, Tremblay Y, Lorentsen S-H, Garthe S, Votier S, Pons JB, Siorat F, Furness RW, Grecian WJ, Bearhop S, Montevecchi WA, Grémillet D (2012): Meta-population evidence of oriented chain-migration in northern gannets. Frontiers in Ecology and the Environment 10: 237-242.
  • Garthe S, Ludynia K, Hüppop O, Kubetzki U, Meraz JF, Furness RW (2012): Energy budgets reveal equal benefits of varied migration strategies in northern gannets. Marine Biology 159: 1907-1915.
  • Garthe S, Markones N, Mendel B, Sonntag N, Krause J (2012): Protected areas for seabirds in German offshore waters: designation, retrospective consideration and current perspectives. Biological Conservation 156: 126-135.
  • Kubetzki U, Garthe S, Dien J (2012): Das Programm "Garden Birds": Die Wintervogelwelt der Gärten in Hamburg und Umgebung. Falke 59: 52-57.
  • Montevecchi WA, Fifield D, Burke C, Garthe S, Hedd A, Rail J-F, Robertson G (2012): Tracking long-distance migration to assess marine pollution impact. Biology Letters 8: 218-221.
  • Montevecchi WA, Hedd A, McFarlane Tranquilla L, Fifield DA, Burke CM, Regular PM, Davoren GK, Garthe S, Robertson GJ, Phillips RA (2012): Tracking seabirds to identify ecologically important and high risk marine areas in the western North Atlantic. Biological Conservation 156: 62-71.
  • Schwemmer P, Güpner F, Guse N, Garthe S (2012): Nahrungswahl von Vogelarten der deutschen Nordseeküste. Vogelwarte 50: 141-154. pdf zum Download (600 KB)
  • Siebert U, Schwemmer P, Guse N, Harder T, Garthe S, Prenger-Berringhof E, Wohlsein P (2012): Health status of seabirds and coastal birds found at the German North Sea coast. Acta Vet Scand 54: 43.
  • Sonntag N, Dries H, Fock H, Bellebaum J, Garthe S (2012): Seabirds, set nets and conservation management: assessment of conflict potential and vulnerability of birds to bycatch in gill nets. ICES Journal of Marine Science 69: 578-589.
  • Stauss C, Bearhop S, Bodey TW, Garthe S, Gunn C, Grecian WJ, Inger R, Knight ME, Newton J, Patrick SC, Phillips RA, Waggitt JJ, Votier SC (2012): Sex-specific foraging behaviour in a monomorphic seabird: Incidence and implications. Marine Ecology Progress Series 457: 151-162.



  • Burkhard B, Opitz S, Lenhart H, Ahrendt K, Garthe S, Mendel B, Windhorst W (2011): Ecosystem based modeling and indication of ecological integrity in the German North Sea - Case study offshore wind parks. Ecological Indicators 11: 168-174.
  • Garthe S, Montevecchi WA, Davoren GK (2011): Inter-annual changes in prey fields trigger different foraging tactics in a large marine predator. Limnology and Oceanography 56: 802-812.
  • Kotzerka J, Hatch SA, Garthe S (2011): Evidence for foraging site fidelity and individual foraging behavior of chick-reaing Pelagic Cormorants in the Gulf of Alaska. Condor 113: 80-88.
  • Kubetzki U, Garthe S (2011): Auswirkungen von Klimaänderungen auf Seevögel. In: Lozán JL, Graßl H, Karbe L, Reise K (Hrsg): Warnsignale Klima - Die Meere: Änderungen und Risiken. Wissenschaftliche Fakten. S. 203-207.
  • Kubetzki U, Garthe S, Hüppop O (2011): Auswirkungen auf See- und Zugvögel: Offshore-Windenergieanlagen. Falke 58: 490-494. pdf zum Download (3 MB)
  • Markones N, Garthe S (2011): Monitoring von Seevögeln im Offshore-Bereich der schleswig-holsteinischen Nordsee im Rahmen von NATURA 2000 in den Jahren 2004 bis 2009. Corax 22: 11-50.
  • Rubarth J, Dreyer A, Guse N, Einax JW, Ebinghaus R (2011): Perfluorinated compounds in red-throated divers from the German Baltic Sea: new findings from their distribution in 10 different tissues. Environmental Chemistry 8: 419-428.
  • Schwemmer P, Colijn F (2011) The Wadden Sea in an international perspective. Current Wadden Sea research in Germany and Denmark. Revised version including Dutch research institutes. A report for the Dutch Waddenacademie.
  • Schwemmer P, Garthe S (2011): Spatial and temporal patterns of habitat use by Eurasian Oystercatchers (Haematopus ostralegus) population in the eastern Wadden Sea as revealed using GPS data loggers. Marine Biology 158: 541-550.
  • Schwemmer P, Mendel B, Sonntag N, Dierschke V, Garthe S (2011): Effects of ship traffic on seabirds in offshore waters: implications for marine conservation and spatial planning. Ecological Applications 21: 1851-1860.
  • Schwemmer P, Tischler T, Rehm R, Garthe S (2011): Habitatnutzung, Verbreitung und Nahrungswahl der Lachmöwe (Larus ridibundus) im küstennahen Binnenland Schleswig-Holsteins. Corax 21: 355-374.
  • Skov H, Heinänen S, Zydelis R, Bellebaum J, Bzoma S, Dagys M, Durinck J, Garthe S, Grishanov G, Hario M, Kieckbusch JJ, Kube J, Kuresoo A, Larsson K, Luigujoe L, Meissner W, Nehls HW, Nilsson L, Petersen IK, Roos MM, Pihl S, Sonntag N, Stock A, Stipniece A, Wahl J (2011): Waterbird populations and pressures in the Baltic Sea. TemaNord 2011:550, Nordic Council of Ministers, Copenhagen. 201 S.
  • van Franeker JA, Blaize C, Danielsen J, Fairclough K, Gollan J, Guse N, Hansen P-L, Heubeck M, Jensen J-K, Le Guillou G, Olsen B, Olsen K-O, Pedersen J, Stienen EWM, Turner DM (2011): Monitoring plastic ingestion by the northern fulmar Fulmarus glacialis in the North Sea. Environmental Pollution 159 : 2609-2615



  • Bellebaum J, Bock C, Garthe S, Kube J, Schilz M, Sonntag N (2010): Vorkommen des Gelbschnabeltauchers Gavia adamsii in der deutschen Ostsee. Vogelwelt 131: 179-184.
  • Burkhard B, Opitz S, Lenhart H, Ahrendt K, Garthe S, Mendel B, Nerge P, Windhorst W (2010): Modellbasierte Bewertung der Auswirkungen von Offshore-Windkraftanlagen auf die ökologische Integrität der Nordsee. In: Kannen A, Schernewski G, Krämer I, Lange M, Janßen H, Stybel N (Hrsg.): Forschung für ein Integriertes Küstenzonenmanagement: Fallbeispiele Odermündungsregion und Offshore-Windkraft in der Nordsee. Coastline Rep. 15: 15-29.
  • Davoren GK, Garthe S, Montevecchi WA, Benvenuti S (2010): Influence of prey behaviour and other predators on the foraging activities of a marine avian predator in a Low Arctic ecosystem. Marine Ecology Progress Series 404: 275-287.
  • Garthe S (2010): Ein Hochseevogel mit spezieller Tauchtechnik: Der Basstölpel. Falke 57: 396-401. pdf zum Download (2 MB)
  • Garthe S, Mendel B (2010): Cumulative effects of anthropogenic activities on seabirds in the North Sea: Methodology and first results. In: Lange M, Burkhard B, Garthe S, Gee K, Kannen A, Lenhart H, Windhorst W (eds): Analyzing coastal and marine changes: Offshore wind farming as a case study. Zukunft Küste - Coastal Futures Synthesis Report. LOICZ Research and Studies 36: 86-89.
  • Garthe S, Mitschke A, Krebs B, Rupnow G (2010): Expansion des Möwen-Brutbestandes im Hamburger Hafen zu Beginn des 21. Jahrhunderts. Hamburger avifaunistische Beiträge 37: 117-142.
  • Hötker H, Schrader S, Schwemmer P, Oberdiek N, Blew J (2010) Status, threats and conservation of birds in the German Wadden Sea. Technical Report NABU.
  • Kotzerka J, Garthe S, Hatch SA (2010): GPS tracking devices reveal foraging strategies of Black-legged Kittiwakes. Journal of Ornithology 151: 459-467.
  • Kubetzki U, Garthe S (2010): Über den Dächern von Kiel und Westerland: Möwen als Dachbrüter in Schleswig-Holstein. Corax 21: 301-309.
  • Ludynia K, Garthe S, Luna-Jorquera G (2010): Distribution and foraging behaviour of the Peruvian Booby (Sula variegata) off northern Chile. Journal of Ornithology 151: 103-111.
  • Ludynia K, Jones R, Kemper J, Garthe S, Underhill LG (2010): Foraging behaviour of Bank Cormorants in Namibia: Implications for conservation. Endangered Species Research 12: 31-40.
  • Markones N, Dierschke V, Garthe S (2010): Seasonal differences in at-sea activity of seabirds underline high energetic demands during the chick-rearing period. Journal of Ornithology 151: 329-336.
  • Markones N, Sonntag N, Garthe S (2010): Seevogelmonitoring in Nord- und Ostsee: Vogelbeobachtung auf offenem Meer. Falke 57: 60-65. pdf zum Download (4 MB)
  • Mendel B, Garthe S (2010): Kumulative Auswirkungen von Offshore-Windkraftnutzung und Schiffsverkehr am Beispiel der Seetaucher in der Deutschen Bucht. In: Kannen A, Schernewski G, Krämer I, Lange M, Janßen H, Stybel N (Hrsg.): Forschung für ein Integriertes Küstenzonenmanagement: Fallbeispiele Odermündungsregion und Offshore-Windkraft in der Nordsee. Coastline Rep. 15: 31-44.
  • Mendel B, Garthe S (2010): Strategien bei der Nahrungssuche: Mit Hightech auf der Spur der Helgoländer Heringsmöwen. Falke 57: 402-408. pdf zum Download (2 MB)
  • Paiva VH, Geraldes P, Ramirez I, Garthe S, Ramos JA (2010): How area restricted search of a pelagic seabird changes while performing a dual foraging strategy. Oikos 119: 1423-1434.
  • Paiva VH, Geraldes P, Ramirez I, Meirinho A, Garthe S, Ramos JA (2010): Foraging plasticity in a pelagic seabird species along a marine productivity gradient. Marine Ecology Progress Series 398: 259-274.
  • Paiva VH, Geraldes P, Ramirez I, Meirinho A, Garthe S, Ramos JA (2010): Oceanographic characteristics of areas used by Cory's shearwaters during short and long foraging trips in the North Atlantic. Marine Biology 157: 1385-1399.
  • Paiva VH, Guilford T, Meade J, Geraldes P, Ramos JA, Garthe S (2010): Flight dynamics of Cory's Shearwater foraging in a coastal environment. Zoology 113: 47-56.
  • Paiva VH, Xavier J, Geraldes P, Ramirez I, Garthe S, Ramos JA (2010): Foraging ecology of Cory's shearwaters in different oceanic environments of the North Atlantic. Marine Ecology Progress Series 410: 257-268.



  • Garthe S , Markones N, Hüppop O, Ader S (2009): Effects of hydrographic and meteorological factors on seasonal seabird abundance in the southern North Sea. Marine Ecology Progress Series 391: 243-255.
  • Garthe S, Schwemmer P, Petersen IK, Laursen K (2009): Offshore Area. Thematic Report No. 17. In: Marencic H, de Vlas J. (Eds.): Quality Status Report 2009. WaddenSea Ecosystem No. 25. Common Wadden Sea Secretariat, Trilateral Monitoring and Assessment Group, Wilhelmshaven, Germany.
  • Guse N, Garthe S, Schirmeister B (2009): Diet of red-throated divers Gavia stellata reflects the seasonal availability of Atlantic herring Clupea harengus in the southern Baltic Sea. Journal of Sea Research 62: 268-275.
  • Hamer KC, Humphreys EM, Magalhaes MC, Garthe S, Hennicke J, Peters G, Grémillet D, Skov H, Wanless S (2009): Fine-scale foraging behaviour of a medium-ranging marine predator. Journal of Animal Ecology 78: 880-889.
  • Kubetzki U, Garthe S, Fifield D, Mendel B, Furness RW (2009): Individual migratory schedules and wintering areas of northern gannets. Marine Ecology Progress Series 391: 257-265.
  • Markones N, Guse N (2009): Eignung von Strandfunden zum Studium der Ernährungsökologie von Basstölpeln (Sula bassana) und Dreizehenmöwen (Rissa tridactyla). Corax 21: 5-12.
  • Markones N, Guse N, Hüppop O, Garthe S (2009): Unchanging diet in a stable colony: Contemporary and past diet composition of Black-legged Kittiwakes Rissa tridactyla at Helgoland, south-eastern North Sea. Helgoland Marine Research 63: 199-206.
  • Montevecchi WA, Benvenuti S, Garthe S, Davoren GK, Fifield D (2009): Flexible foraging tactics by a large opportunistic seabird preying on forage and large pelagic fishes. Marine Ecology Progress Series 385: 295-306.
  • Schwemmer P, Guse N, Markones N, Adler S, Garthe S (2009): Influence of water flow velocity and water depth on distribution and foraging patterns of a surface-feeding seabird in the German Wadden Sea. Fisheries Oceanography 18: 161-172.
  • Sonntag N, Garthe S, Adler S (2009): A freshwater species wintering in a brackish environment: Habitat selection and diet of Slavonian grebes (Podiceps auritus) in the southern Baltic Sea. Estuarine and Coastal Shelf Science 84: 186-194.
  • Zydelis R, Bellebaum J, Österblom H, Vetemaa M, Schirmeister B, Stipniece A, Dagys M, van Eerden M, Garthe S (2009): Bycatch in gillnet fisheries - An overlooked threat to waterbird populations. Biological Conservation 142: 1269-1281.



  • Garthe S, Guse N, Sonntag N (2008): Spatio-temporal patterns of inshore and offshore foraging by Great Cormorants in the southwestern Baltic Sea. In: Wollny-Goerke K, Eskildsen K (eds): Marine mammals and seabirds in front of offshore wind energy. MINOS - Marine warm-blooded animals in North and Baltic Seas. Teubner, Wiesbaden. pp. 143-153.
  • Garthe S, Schwemmer P (2008): Durchzug, Sommer- und Wintervorkommen der Zwergmöwe Hydrocoloeus minutus an der Unterelbe. Vogelkundliche Berichte Niedersachsen 40: 399-407.
  • Markones N, Garthe S, Dierschke V, Adler S (2008): Small scale temporal variability of seabird distribution patterns in the south-eastern North Sea. In: Wollny-Goerke K, Eskildsen K (eds): Marine mammals and seabirds in front of offshore wind energy. MINOS - Marine warm-blooded animals in North and Baltic Seas. Teubner, Wiesbaden. pp. 115-140.
  • Mendel B, Sonntag N, Wahl J, Schwemmer P, Dries H, Guse N, Müller S, Garthe S (2008): Artensteckbriefe von See- und Wasservögeln der deutschen Nord- und Ostsee. Verbreitung, Ökologie und Empfindlichkeiten gegenüber Eingriffen in ihren marinen Lebensraum. Naturschutz und Biologische Vielfalt 59. Bundesamt für Naturschutz, Bonn - Bad Godesberg.
  • Mendel B, Sonntag N, Wahl J, Schwemmer P, Dries H, Guse N, Müller S, Garthe S (2008): Profiles of seabirds and waterbirds of the German North and Baltic Seas. Distribution, ecology and sensitivities to human activities within the marine enviroment. Naturschutz und Biologische Vielfalt 61. Bundesamt für Naturschutz, Bonn - Bad Godesberg.
  • Schwemmer P, Garthe S, Mundry R (2008): Area utilization of gulls in a coastal farmland landscape: habitat mosaic supports niche segregation of opportunistic species. Landscape Ecology 23: 355-367.
  • Schwemmer P, Garthe S (2008): Regular habitat switch as an important feeding strategy of an opportunistic seabird species at the interface between land and sea. Estuarine, Coastal and Shelf Science 77: 12-22.
  • Skov H, Humphreys E, Garthe S, Geitner K, Grémillet D, Hamer K, Hennicke J, Parner H, Wanless S (2008): Application of spatial modelling to GPS tracking data of marine mammals as a means for analyzing their feeding habits. Ecological Modelling 212: 504-512.
  • Wilson RP, FH Vargas, A Steinfurth, P Riordan, Y Ropert-Coudert & DW Macdonald (2008): What grounds some birds for life? Movement and diving in the sexually dimorphic Galápagos Cormorant. Ecological Monographs 78: 633-652.



  • Barrett, RT, Camphuysen CJ, Anker-Nilssen T, Chardine JW, Furness RW, Garthe S, Hüppop O, Leopold MF, Montevecchi WA, Veit RR (2007): Diet studies of seabirds: a review and recommendations. ICES Journal of Marine Science 64: 1675-1691.
  • Davoren GK, May C, Penton P, Reinfort B, Buren A , Burke C, Andrews D, Montevecchi WA, Record N, deYoung B, Rose-Taylor C, Bell T, Anderson JT, Koen-Alonso M, Garthe S (2007): An ecosystem-based research program for capelin (Mallotus villosus) in the Northwest Atlantic: overview and results. Journal of the Northwest Atlantic Fisheries Science 39: 35-48.
  • Ehrich S, Adlerstein S, Brockmann U, Floeter J, Garthe S, Hinz H, Kröncke I, Neumann H, Reiss H, Sell AF, Stein M, Stelzenmüller V, Stransky C, Temming A, Wegner G, Zauke G-P (2007): 20 years of the German Small-Scale Bottom Trawl Survey (GSBTS): A review. Senckenbergiana maritima 37: 13-82.
  • Garthe S , Flore B-O (2007): Population trend over 100 years and conservation needs of breeding sandwich terns (Sterna sandvicensis) at the German North Sea coast. Journal of Ornithology 148: 215-227.
  • Garthe S , Montevecchi WA, Chapdelaine G, Rail J-F, Hedd A (2007): Contrasting foraging tactics by northern gannets (Sula bassana) breeding in different oceanographic domains with different prey fields. Marine Biology 151: 687-694.
  • Garthe S , Montevecchi WA, Davoren G (2007): Flight destinations and foraging behaviour of northern gannets (Sula bassana) preying on a small foraging fish in a Low Arctic ecosystem. Deep-Sea Research Part II 54: 311-320.
  • Garthe S, Sonntag N, Schwemmer P, Dierschke V (2007): Estimation of seabird numbers in the German North Sea throughout the annual cycle and their biogeographic importance. Vogelwelt 128: 163-178.
  • Hamer KC, Humphreys EM, Garthe S, Hennicke J, Peters G, Phillips RA, Harris MP, Wanless S (2007): Annual variations in diet, feeding locations and foraging behaviour of gannets in the North Sea: flexibility, consistency and contraint. Marine Ecology Progress Series 338: 295-305.
  • Kubetzki U, Garthe S (2007): Nests with a view: Distribution, nest habitats and diets of roof-breeding Common Gulls (Larus canus) in northern Germany. Waterbirds 30: 602-608.
  • Markones N, Guse N (2007): Räumlich-zeitliche Verteilung und Nahrungserwerbsstrategien von Silbermöwen Larus argentatus der westlichen Ostsee: Erkenntnisse einer Ringfundanalyse.Vogelwarte 45: 1-13. pdf zum Download (2 MB)
  • Mendel B, Sonntag N, Garthe S (2007): GIS-gestützte Analyse zur Verbreitung und Habitatwahl ausgewählter Seevogelarten in der Ostsee. In: Traub K-P, Kohlus J (Hrsg.): Geoinformationen für die Küstenzone. Beiträge des 1. Hamburger Symposiums zur Küstenzone. Herbert Wichmann Verlag, Heidelberg. S. 196-206.
  • Villablanca R, Luna-Jorquera G, Marín VH, Garthe S, Simeone A (2007): How does a generalist seabird species use its marine habitat? The case of the Kelp Gull in a coastal upwelling area of the Humboldt Current. ICES Journal of Marine Science 64: 1348-1355.



  • Bellebaum J, Garthe S, Kube J, Nehls HW, Schulz A, Skov H (2006): Wasservögel im Küstenmeer Mecklenburg-Vorpommerns: ein Überblick zu Bestandssituation, Gefährdungen und Abgrenzungen neuer Vogelschutzgebiete. Berichte zum Vogelschutz 43: 31-47.
  • Colijn F, Garthe S (2006): EU Directives and their effect on the ecosystem of the Wadden Sea. In Laursen K (ed): Monitoring and assessment in the Wadden Sea. Proceedings from the 11th International Scientific Wadden Sea Symposium, Esbjerg, Denmark. NERI Technical Report 573: 13-20.
  • Dierschke V, Garthe S. (2006): Literature review of offshore wind farms with regards to seabirds. BfN-Skripten 186: 131-198.
  • Dierschke V, Lorentzen NH (2006): Phänologie südnorwegischer Mantelmöwen Larus marinus auf Helgoland (Deutsche Bucht). Vogelwelt 127: 31-36.
  • Dierschke V, Garthe S, Mendel B (2006): Possible conflicts between offshore wind farms and seabirds in the German sectors of North Sea and Baltic Sea. In: Köller J, Köppel H, Peters W (eds): Offshore wind energy. Research on environmental impacts. Springer. pp. 121-143.
  • Furness RW, Crane JE, Bearhop S, Garthe S, Käkelä A, Käkelä R, Kelly A, Kubetzki U, Votier SC, Waldron S (2006): Techniques to link migration patterns of seabirds with diet specialization, condition and breeding performance. Ardea 94: 631-638.
  • Garthe S (2006): Identification of areas of seabird concentrations in the German North Sea and Baltic Sea using aerial and ship-based surveys: In: von Nordheim, H., Boedeker, D. & Krause, J.C. (eds.): Progress in marine conservation in Europe: Natura 2000 sites in German offshore waters. Springer. pp. 225-238.
  • Garthe S, Skov H (2006): Selection of suitable sites for marine protected areas for seabirds: a case study with Special Protection Areas (SPAs) in the German Baltic Sea. In: Boere GC, Galbraith CA, Stroud DA (eds): Waterbirds around the world. The Stationary Office, Edinburgh, UK. pp. 739-742.
  • Goss-Custard, JD, AD West, MG Yates, RWG Caldow, RA Stillman, L Bardsley, J Castilla, M Castro, V Dierschke, SEA Le V dit Durell, G Eichhorn, BJ Ens, K-M Exo, PU Udayangani-Fernando, PN Ferns, PAR Hockey, JA Gill, I Johnstone, B Kalejta-Summers, JA Masero, F Moreira, R Velu Nagarajan, IPF Owens, C Pacheco, A Perez-Hurtado, D Rogers, G Scheiffarth, H Sitters, WJ Sutherland, P Triplet, DH Worrall1, Y Zharikov, L Zwarts & RA Pettifor (2006): Intake rates and the functional response in shorebirds (Charadriiformes) eating macro-invertebrates. Cambridge Philosophical Society, Biol. Rev. 81: 501-529.
  • Hamer KC, Lewis S, Wanless S, Phillips RA, Sherratt TN, Humphreys EM, Hennicke J, Garthe S (2006): Use of gannets to monitor prey availability in the NE Atlantic Ocean: colony size, diet and foraging behaviour. In: Boyd IL, Wanless S, Camphuysen CJ (eds): Top predators in marine ecosystems. Their role in monitoring and management. Cambridge University Press, Cambridge. pp. 236-248.
  • Montevecchi WA, Garthe S, Davoren GK (2006): Biophysical influences on seabird trophic assessments. In: Boyd IL, Wanless S, Camphuysen CJ (eds): Top predators in marine ecosystems. Their role in monitoring and management. Cambridge University Press, Cambridge. pp. 118-130.
  • Schwemmer P, Garthe S (2006): Spatial patterns in at-sea behaviour during spring migration by Little Gulls (Larus minutus) in the south-eastern North Sea. Journal of Ornithology 147: 354-366.
  • Sonntag N, Mendel B, Garthe S (2006): Die Verbreitung von See- und Wasservögeln in der deutschen Ostsee im Jahresverlauf. Vogelwarte 44: 81-112. pdf zum Download (3 MB)
  • Sonntag N, Weichler T, Weiel S, Meyer B (2006): Blinder Passagier - Zweifarbfledermaus (Vespertilio murinus) landet auf einem Forschungsschiff in der Pommerschen Bucht (südliche Ostsee). Nyctalus 11: 277-279.



  • Dierschke V, B Mendel & H Schmaljohann (2005): Differentidal timing of spring migration in northern wheatears Oenanthe oenanthe: hurried males or weak females? Behav. Ecol. Sociobiol. 57: 470-480.
  • Froese R, Garthe S, Pauly D, Piatkowski U (2005): Trophic signatures of marine organisms in the Mediterranean as compared with other ecosystems. Belgian Journal of Zoology 135: 139-143.
  • Garthe S , Schwemmer P (2005): Seabirds at Sea-Untersuchungen in den deutschen Meeresgebieten. Vogelwelt 126: 67-74.
  • Guse N, Fleet D, van Franeker J, Garthe S (2005): Der Eissturmvogel (Fulmarus glacialis) – Mülleimer der Nordsee? Seevögel 26 (2): 3-12.
  • Ludynia K, Garthe S, Luna-Jorquera G (2005): Seasonal and regional variation in the diet of the Kelp Gull in northern Chile. Waterbirds 28: 359-365.
  • Schwemmer P, Garthe S (2005): At-sea distribution and behaviour of a surface-feeding seabird, the lesser black-backed gull Larus fuscus, and its association with different prey. Marine Ecology Progress Series 285: 245-258.



  • Camphuysen CJ, Garthe S (2004): Recording foraging seabirds at sea. Standardised recording and coding of foraging behaviour and multi-species foraging associations. Atlantic Seabirds 6: 1-32.
  • Dierschke V, Garthe S, Markones N (2004): Aktionsradien Helgoländer Dreizehenmöwen Rissa tridactyla und Trottellummen Uria aalge während der Aufzuchtphase. Vogelwelt 125: 11-19.
  • Exo K-M, Hüppop O & Garthe S (2004): Offshore wind farms and bird conservation: potential conflicts and minimum requirements for project-related studies in the North Sea and the Baltic Sea. In: Ferrer M., de Lucas M., Janss G. (eds.): Birds and wind power. Quercus, Spain.
  • Garthe S , Hüppop O (2004): Scaling possible adverse effects of marine wind farms on seabirds: developing and applying a vulnerability index. Journal of Applied Ecology 41: 724-734.
  • Garthe S , Montevecchi WA, Ojowski U, Stenhouse IJ (2004): Diets of northern fulmar (Fulmarus glacialis) chicks in the north-west Atlantic. Polar Biology 27: 277-280.
  • Sonntag N, Engelhard O, Garthe S (2004): Sommer- und Mauservorkommen von Trauer- und Samtenten (Melanitta nigra und M. fusca) auf der Oderbank. Vogelwelt 125: 77-82.
  • Weichler T, Garthe S, Luna-Jorquera G, Moraga J (2004): Seabird distribution on the Humboldt Current in northern Chile in relation to hydrography, productivity, and fisheries. ICES Journal of Marine Science 61: 148-154.
  • Wilson RP, A Scolaro, F Quintana, U Siebert, MT Straten, K Mills, I Zimmer, N Liebsch, A Steinfurth, G Spindler & G Müller (2004): To the bottom of the heart: Cloacal movement as an index of cardiac frequency, respiration and digestive evacuation in penguins. Marine Biology 144: 813-827.



  • Dierschke V, Hüppop O, Garthe S (2003): Populationsbiologische Schwellen der Unzulässigkeit für Beeinträchtigungen der Meeresumwelt am Beispiel der in der deutschen Nord- und Ostsee vorkommenden Vogelarten. Seevögel 24: 61-72.
  • Exo, K-M, Hälterlein B, Blew J, Garthe S, Hüppop O, Südbeck P, Scheiffarth G (2003): Küsten- und Seevögel. In: Lozán JL, Rachor E, Reise K, Sündermann J, von Westernhagen H (Hrsg): Warnsignale aus Nordsee & Wattenmeer. Eine aktuelle Umweltbilanz. Wissenschaftliche Auswertungen, Hamburg. pp. 317-329.
  • Exo K-M, Hüppop O, Garthe S (2003): Birds and offshore wind farms: a hot topic in marine ecology. Wader Study Group Bulletin 100: 50-53.
  • Garthe S (2003) : Verbreitung, Bestand und Jahresdynamik der Mantelmöwe Larus marinus in der Deutschen Bucht, Nordsee. Corax 19, Sonderheft 2: 43-50.
  • Garthe S (2003): Verteilungsmuster und Bestände von Seevögeln in der Ausschließlichen Wirtschaftszone (AWZ) der deutschen Nord- und Ostsee und Fachvorschläge für EU-Vogelschutzgebiete. Berichte zum Vogelschutz 40: 15-56.
  • Garthe S, Benvenuti S, Montevecchi WA (2003): Temporal patterns of foraging activities of northern gannets Morus bassanus in the north-west Atlantic. Canadian Journal of Zoology 81: 453-461.
  • Garthe S, Krüger T, Kubetzki U, Weichler T (2003): Monitoring von Seevögeln auf See: Gegenwärtiger Stand und Perspektiven. In: Anon.: Vogelmonitoring in Deutschland. Berichte des Landesamtes für Umweltschutz Sachsen-Anhalt, Sonderheft 1/2003: 62-64.
  • Garthe S , Scherp B (2003): Utilization of discards and offal from commercial fisheries by seabirds in the Baltic Sea. ICES Journal of Marine Science 60: 980-989.
  • Garthe S, Ullrich N, Weichler T, Dierschke V, Kubetzki U, Kotzerka J, Krüger T, Sonntag N, Helbig AJ (2003): See- und Wasservögel der deutschen Ostsee - Verbreitung, Gefährdung und Schutz. Bundesamt für Naturschutz, Bonn. 170 pp.
  • Garthe S, Weichler T, Krüger T (2003): Seabirds at Sea-Untersuchungen in Nord- und Ostsee: von den Grundlagen bis zur Anwendung. Corax 19, Sonderheft 2: 1-6.
  • Garthe S, Wienck K, Cassens I (2003): Herring Gull Larus argentatus winter diet at the western Baltic Sea coast: does ice cover make a difference? Atlantic Seabirds 5: 13-20.
  • Krüger T, Garthe S (2003): Truppgröße und Truppzusammensetzung von See- und Küstenvögeln auf dem Wegzug vor Wangerooge. Corax 19, Sonderheft 2: 7-14.
  • Kubetzki U, Garthe S (2003): Distribution, diet and habitat selection by four sympatrically breeding gull species in the southeastern North Sea. Marine Biology 143: 199-207.
  • Simeone A, Luna-Jorquera G, Bernal M, Garthe S, Sepúlveda F, Villablanca R, Ellenberg U, Contreras M, Muñoz J, Ponce T (2003): Breeding distribution and abundance of seabirds on islands off north-central Chile. Revista Chilena de Historia Natural 76: 323-333.
  • Tasker ML, Camphuysen K, Garthe S, Anker-Nilssen T (2003): The alteration of the composition of seabird communities by fisheries. In: Tasker ML, Furness RW (eds): Seabirds as monitors of the marine environment. ICES Cooperative Research Report 258: 37-51.
  • Wahl J, Blew J, Garthe S, Günther K, Mooij J, Sudfeldt C (2003): Überwinternde Wasser- und Watvögel in Deutschland: Bestandsgrößen und Trends ausgewählter Vogelarten für den Zeitraum 1990-2000. Berichte zum Vogelschutz 40: 91-103.



  • Exo K-M, Hüppop O, Garthe S (2002): Offshore-Windenergieanlagen und Vogelschutz. Seevögel 23: 83-95.
  • Garthe S , Hüppop O, Weichler T (2002): Anleitung zur Erfassung von Seevögeln auf See von Schiffen. Seevögel 23: 47-55.
  • Hüppop O, Exo K-M, Garthe S (2002): Empfehlungen für projektbezogene Untersuchungen möglicher bau- und betriebsbedingter Auswirkungen von Offshore-Windenergieanlagen auf Vögel. Berichte zum Vogelschutz 39: 75-94.
  • Krüger T, Garthe S (2002): Vorkommen ausgewählter See- und Küstenvögel vor Wangerooge: Der Einfluss von Windrichtung und -stärke. Journal für Ornithologie 143: 155-170.
  • Simeone A, Garthe S, Sepúlveda F, Luna-Jorquera G (2002): Sula nebouxii en Isla Pájaros, Región de Coquimbo. Boletín Chileno de Ornitología 9: 48.
  • Wilson RP, Grémillet D, Syder J, Kierspel MAM, Garthe S, Weimerskirch H, Schäfer-Neth C, Scolaro JA, Bost C-A, Plötz J, Nel D (2002): Remote-sensing systems and seabirds: their use, abuse and potential for measuring marine environmental variables. Marine Ecology Progress Series 228: 241-261.



  • Froese R, Piatkowski U, Garthe S, Pauly D (2001): A preliminary comparison of the trophic structure of some large marine ecosystems. ICES Doc. CM 2001/T:07.
  • Froese R, Piatkowski U, Garthe S, Pauly D (2001): Comparing the North Sea with the Caribbean? ICES Newsletter 38: 24-25.
  • Garthe S , Furness RW (2001): Frequent shallow diving by a Northern Fulmar feeding at Shetland. Waterbirds 24: 287-289.
  • Krüger T, Garthe S (2001): Flight altitudes of coastal birds in relation to wind direction and speed. Atlantic Seabirds 3: 203-216.
  • Krüger T, Garthe S (2001): Tagesperiodik von See- und Küstenvögeln auf dem Wegzug vor Wangerooge. Vogelkundliche Berichte Niedersachsen 33: 25-34.
  • Mitschke A, Garthe S , Hüppop O (2001): Erfassung der Verbreitung, Häufigkeiten und Wanderungen von See- und Wasservögeln in der deutschen Nordsee und Entwicklung eines Konzeptes zur Umsetzung internationaler Naturschutzziele. BfN-Skripten 34: 1-100.
  • Ojowski U, Eidtmann C, Furness RW, Garthe S (2001): Diet and nest attendance of incubating and chick-rearing Northern Fulmars (Fulmarus glacialis) in Shetland. Marine Biology 139: 1193-1200.



  • Garthe S (2000): Mögliche Auswirkungen von Offshore-Windenergieanlagen auf See- und Wasservögel der deutschen Nord- und Ostsee. In: Merck T, von Nordheim H (eds): Technische Eingriffe in marine Lebensräume. Workshop des Bundesamtes für Naturschutz, Internationale Naturschutzakademie Insel Vilm, 27.-29. Oktober 1999. BfN-Skripten 29: 113-119.
  • Garthe S (2000): Von Brutvögeln und Durchzüglern. In: Newig J, Theede H (eds): Pflanzen- und Tierwelt im Wattenmeer. Ellert & Richter, Hamburg. S. 100-117.
  • Garthe S, Benvenuti S, Montevecchi WA (2000): Pursuit-plunging by Northern Gannets (Sula bassana) feeding on Capelin (Mallotus villosus). Proceedings of the Royal Society London, Biological Sciences 267: 1717-1722.
  • Garthe S, Flore B-O, Hälterlein B, Hüppop O, Kubetzki U, Südbeck P (2000): Die Brutbestandsentwicklung der Möwen (Laridae) an der deutschen Nordseeküste in der zweiten Hälfte des 20. Jahrhunderts. Vogelwelt 121: 1-13.
  • Garthe S, Hüppop O (2000): Aktuelle Entwicklungen beim Seabirds-at-Sea-Programm in Deutschland. Vogelwelt 121: 301-305.
  • Luna-Jorquera G, Garthe S, Sepúlveda FG, Weichler T, Vásquez JA (2000): Population size of Humboldt Penguins assessed by combined terrestrial and at-sea counts. Waterbirds 23: 506-510.
  • Mitschke A, Garthe S, Mulsow R (2000): Langfristige Bestandstrends von häufigen Brutvögeln in Hamburg. Vogelwelt 121: 155-164.
  • Tasker ML, Camphuysen CJ, Cooper J, Garthe S, Montevecchi WA, Blaber SJM (2000): The impacts of fishing on marine birds. ICES Journal of Marine Science 57: 531-547.



  • Garthe S (1999): The influence of fishing activities on the distribution and feeding ecology of seabirds at sea. In: Adams NJ, Slotow RH (eds): Proc. 22 Int. Ornithol. Congr., Durban: 706-716. BirdLife South Africa, Johannesburg.
  • Garthe S, Freyer T, Hüppop O, Wölke D (1999): Breeding Lesser Black-backed Gulls Larus graellsii and Herring Gulls Larus argentatus: coexistence or competition? Ardea 87: 227-236.
  • Garthe S, Grémillet D, Furness RW (1999): At-sea-activity and foraging efficiency in chick-rearing Northern Gannets (Sula bassana): a case study in Shetland. Marine Ecology Progress Series 185: 93-99.
  • Garthe S, Hüppop O (1999): Effect of ship speed on seabird counts in areas supporting commercial fisheries. Journal of Field Ornithology 70: 28-32.
  • Garthe S, Kubetzki U, Hüppop O, Freyer T (1999): Zur Ernährungsökologie von Herings-, Silber- und Sturmmöwe (Larus fuscus, L. argentatus und L. canus) auf der Nordseeinsel Amrum während der Brutzeit. Seevögel 20: 52-58.
  • Garthe S, Walter U, Tasker ML, Becker PH, Chapdelaine G, Furness RW (1999): Evaluation of the role of discards in supporting bird populations and their effects on the species composition of seabirds in the North Sea. In: Furness RW, Tasker ML (eds): Diets of seabirds and consequences of changes in food supply. ICES Cooperative Research Report 232: 29-41.
  • Garthe S, Weimerskirch H (1999): Benefits and threats from fisheries to marine birds. In: Adams NJ, Slotow RH (eds): Proc. 22 Int. Ornithol. Congr., Durban: 647. BirdLife South Africa, Johannesburg.
  • Kubetzki U, Garthe S, Hüppop O (1999): The diet of Common Gulls (Larus canus) breeding on the German North Sea coast. Atlantic Seabirds 1: 57-70.



  • Garthe S (1998): Gleich und doch anders: Zur Habitatwahl von Eissturmvogel (Fulmarus glacialis) und Sturmmöwe (Larus canus) in der Deutschen Bucht. Seevögel 19, Sonderheft: 81-85.
  • Garthe S, Hüppop O (1998): Foraging success, kleptoparasitism and feeding techniques in scavenging seabirds; does crime pay? Helgoländer Meeresuntersuchungen 52: 187-196.
  • Garthe S, Hüppop O (1998): Possible bias in experiments evaluating the consumption of discards by seabirds in the North Sea. Marine Biology 131: 735-741.
  • Garthe S, Kubetzki U (1998): Diet of Sandwich Terns (Sterna sandvicensis) on Juist. Sula 12: 13-18.



  • Camphuysen CJ, Garthe S (1997): An evaluation of the distribution and scavenging habits of Northern Fulmars Fulmarus glacialis in the North Sea. ICES Journal of Marine Science 54: 654-683.
  • Garthe S (1997): Influence of hydrography, fishing activity and colony location on summer seabird distribution in the southeastern North Sea. ICES Journal of Marine Science 54: 566-577.
  • Garthe S, Damm U (1997): Discards in beam trawl fisheries in the German Bight. Archive of Fishery and Marine Research 45: 223-242.
  • Garthe S, Hüppop O (1997): Can seabirds be used as hydrocasts? In: Extended abstracts, Symposium "New Challenges for North Sea Research - 20 years after FLEX '76". Berichte des Zentrums für Meeres- und Klimaforschung in Hamburg, Reihe Z, 2: 77-81.



  • Flore B-O, Garthe S, Degen A (1996): Past and present occurrence of Little Auks Alle alle in Germany. Sula 10: 183-192.
  • Garthe S (Hrsg, 1996): Die Vogelwelt von Hamburg und Umgebung. Band 3. Wachholtz Verlag, Neumünster. 496 S.
  • Garthe S, Bruster K-H (1996): Zunahme von Dachbruten des Austernfischers (Haematopus ostralegus) in Hamburg und Umgebung. Hamburger avifaunistische Beiträge 28: 159-163.
  • Garthe S, Camphuysen CJ, Furness RW (1996): Amounts of discards by commercial fisheries and their significance as food for seabirds in the North Sea. Marine Ecology Progress Series 136: 1-11.
  • Garthe S, Hüppop O (1996): Das "Seabirds-at-sea"-Programm. Vogelwelt 117: 303-305.
  • Garthe S, Hüppop O (1996): Nocturnal scavenging by gulls in the southern North Sea. Colonial Waterbirds 19: 232-241.
  • Garthe S, Ludwig J, Becker PH (1996): Gefährdung der Vogelwelt an Flüssen. In: Lozán JL, Kausch H (eds): Warnsignale aus Flüssen und Ästuaren. Wissenschaftliche Fakten. Blackwell Wissenschaft, Berlin. S. 234-240.
  • Garthe S, Mitschke A (1996): Das Ramsar-Gebiet "Mühlenberger Loch" - ein Süßwasserwatt im Elbästuar. 30jährige Entwicklung der Avifauna und Naturschutz-Bewertung. Naturschutz und Landschaftsplanung 28: 75-80.



  • Camphuysen CJ, Calvo B, Durinck J, Ensor K, Follestad A, Furness RW, Garthe S, Leaper G, Skov H, Tasker ML, Winter CJN (1995) Consumption of discards by seabirds in the North Sea. Final report EC DG XIV research contract BIOECO/93/10. NIOZ-rapport 1995-5, Netherlands Institute for Sea Research, Texel
  • Garthe S, Alicki K, Hüppop O, Sprotte B (1995): Die Verbreitung und Häufigkeit ausgewählter See- und Küstenvogelarten während der Brutzeit in der südöstlichen Nordsee. Journal für Ornithologie 136: 253-266.
  • Hüppop O, Garthe S (1995): Seevögel und Fischerei. Ein Situationsbericht mit Schwerpunkt Nordsee. In Bezzel E (eds): Ornithologen-Kalender '96. Jahrbuch für Vogelkunde und Vogelschutz. Aula-Verlag, Wiesbaden. S. 185-196.
  • Hüppop O, Garthe S (1995): Vögel und Fischerei in der Nordsee. Deutsche Hydrographische Zeitschrift, Supplement 2: 109-115.
  • Leopold M, van Damme C, Garthe S (1995): Grote concentraties Roodkeelduikers Gavia stellata tussen Cuxhaven en Helgoland. Sula 9: 75-78.



  • Garthe S, Hüppop O (1994): Distribution of ship-following seabirds and their utilization of discards in the North Sea in summer. Marine Ecology Progress Series 106: 1-9.
  • Garthe S, Mitschke A (1994): Artenhilfsprogramm und Rote Liste der gefährdeten Brutvögel in Hamburg. Naturschutz und Landschaftspflege in Hamburg, Band 41. 160 pp.
  • Hüppop O, Garthe S (1994): Vögel und Fischerei in der Nordsee. Arbeiten des Deutschen Fischerei-Verbandes e.V., Heft 60: 112-124.
  • Hüppop O, Garthe S, Hartwig E, Walter U (1994): Fischerei und Schiffsverkehr: Vorteil oder Problem für See- und Küstenvögel? In: Lozan JL, Rachor E, Reise K, von Westernhagen H, Lenz W (Hrsg): Warnsignale aus dem Wattenmeer. Wissenschaftliche Fakten. Blackwell Wissenschafts-Verlag, Berlin. pp. 278-285.
  • Mitschke A, Baumung S, Dörnbach R, Garthe S, Klemp S, Schlorf D, Schumacher H-U (1994): Ornithologischer Jahresbericht 1987 bis 1990 für das Hamburger Berichtsgebiet. Hamburger avifaunistische Beiträge 26: 7-97.
  • Mitschke A, Garthe S (1994): Die Bedeutung des Mühlenberger Loches als Rast- und Nahrungsgebiet für Wasser- und Watvögel. Hamburger avifaunistische Beiträge 26: 99-235.
  • Walter U, Hüppop O, Garthe S (1994): Eine komplexe Dreiecksbeziehung - Seevögel, Fischbestände und Fischerei. Schriftenreihe der Schutzgemeinschaft Deutsche Nordseeküste e.V., Band 8: 80-108.



  • Camphuysen CJ, Ensor K, Furness RW, Garthe S, Hüppop O, Leaper G, Offringa H, Tasker ML (1993): Seabirds feeding on discards in winter in the North Sea. EC DG XIV research contract 92/3505. NIOZ-rapport 1993-8. Netherlands Institute for Sea Research, Texel. 142 pp.
  • Garthe S  (1993): Durchzug und Wintervorkommen der Zwergmöwe (Larus minutus) bei Helgoland in den Jahren 1977 bis 1991. Die Vogelwarte 37: 118-129.
  • Garthe S (1993): Möwen und Seeschwalben auf der Unterelbe zwischen Hamburg und Pagensand 1987 bis 1991. Corax 15: 261-269.
  • Garthe S (1993): Quantifizierung von Abfall und Beifang der Fischerei in der südöstlichen Nordsee und deren Nutzung durch Seevögel. Hamburger avifaunistische Beiträge 25: 125-237.
  • Garthe S, Hüppop O (1993): Gulls and Fulmars following ships and feeding on discards at night. Ornis Svecica 3: 159-161.
  • Garthe S, Hüppop O (1993): Offshore studies by the Institute for Bird Study on Helgoland. Seabird Group Newsletter 66: 6-8.
  • Hüppop O, Garthe S (1993): Seabirds and fisheries in the southeastern North Sea. Sula 7: 9-14.



  • Garthe S (1990): Das Vorkommen des Kolkraben (Corvus corax) im Hamburger Raum. Hamburger avifaunistische Beiträge 22: 1-23.



  • Garthe S  (1988): Ungewöhnlicher Brutplatz der Bachstelze (Motacilla alba). Hamburger avifaunistische Beiträge 21: 191-192.