Publications Group Heubel


  • Fortes Silva R, Simon M-V, Heubel K, Borcherding R (2019): Have a break or keep going - behavioral and metabolic overwintering strategies of Neogobius melanostomus and Neogobius fluviatilis, two invasive species of the River Rhine, Germany. Environmental Biology of Fishes
  • Pärssinen V, Kalb N, Vallon M, Anthes N, Heubel K (2019): Male and female preferences for nest characteristics under paternal care. Ecology and Evolution 00:1-12
  • Borcherding J, Heubel K, Storm S (2019): Competition fluctuates across years and seasons in a 6-species-fish community: empirical evidence from the field. Rev Fish Biol Fisheries,


  • de Jong K, Amorim MCP, Fonseca PJ, Heubel KU (2018) Noise affects multimodal communication during courtship in a marine fish. Frontiers in Ecology and Evolution 6(113).
  • de Jong K, Amorim MCP, Fonseca PJ, Klein A, Fox CJ, Heubel KU (2018): Noise can affect acoustic communication and subsequent spawning success in fish. Environmental Pollution 237:814-823.
  • Heubel K (2018): Female mating competition alters female mating preferences in common gobies (Pomatoschistus microps). Current Zoology 64(3):351-361.
  • Mück IM, Heubel, K (2018): Ecological variation along the salinity gradient in the Baltic Sea and its consequences for mating success in the common goby. Current Zoology 64(2):259-270.


  • de Jong K, Schulte G, Heubel KU (2017): The noise egg: A cheap and simple device to produce low-frequency underwater noise for laboratory and field experiments. Methods in Ecology and Evolution 8:268–274.
  • Vallon M & Heubel KU (2017): Egg density and salinity influence filial cannibalism in common gobies. Behavioral Ecology and Sociobiology 71:159.


  • Blom EL, Mück I, Heubel K, and Svensson O (2016): Acoustic and visual courtship traits in two sympatric marine Gobiidae species - Pomatoschistus microps and Pomatoschistus minutus", Environmental Biology of Fishes 99(12): 999-1007.
  • de Jong K, Amorim MCP, Fonseca PJ, Klein A, Heubel KU (2016): Noise affects acoustic courtship behavior similarly in two species of gobies. Proceedings of Meetings on Acoustics 27: 010018; 1-10.
  • Kalb N, Lindström K, Sprenger D, Anthes N, Heubel KU (2016): Male personality and female spawning consistency in a goby with exclusive male care. Behavioral Ecology and Sociobiology 70(5): 683-693.
  • Vallon M, Anthes N, Heubel KU (2016): Water mold infection but not paternity induces selective filial cannibalism in a goby. Ecology and Evolution 6(20):7221-7229.
  • Vallon M, Heubel KU (2016): Old but gold - Males preferentially cannibalize young eggs. Behavioral Ecology and Sociobiology 70(4): 569-573.
  • Vallon M, Grom C, Kalb N, Sprenger N, Anthes N, Lindström K, Heubel KU (2016): You eat what you are: Personality-dependent filial cannibalism in a fish with paternal care. Ecology and Evolution 6(5): 1340-1352.

Earlier Publications

  • Lehtonen J, Schmidt DJ, Heubel K, Kokko, H. (2013): Evolutionary and ecological implications of sexual parasitism. Trends in Ecology and Evolution 28:297-306.
  • Kokko, H. and Heubel, KU (2013): On depending on fish for a living, and other difficulties of living sustainably, in: K. Sterelny, R. Joyce, B. Calcott, and B. Fraser (Eds.), Cooperation and its Evolution Book Series, pages 75-88. MIT press.
  • Kokko H., Heubel KU. (2011): Prudent males, group adaptation, and the tragedy of the commons. Oikos 120(5):641-656.
  • Heubel KU, Rankin DJ, Kokko H (2009): How to go extinct by mating too much: Population consequences of male mate choice and efficiency in a sexual-asexual species complex. Oikos 118(4):513-520.
  • Heubel KU, Schlupp I (2008): Seasonal plasticity in male mating preferences in sailfin mollies. Behavioral Ecology 19(6):1080-1086.
  • Heubel KU, Plath M (2008): Influence of male harassment and female competition on female feeding behaviour in a sexual-asexual mating complex of mollies (Poecilia mexicana, P. formosa). Behavioral Ecology and Sociobiology 62(11):1689-1699.
  • Kokko H, Heubel K (2008): Condition-dependence, genotype-by-environment interactions and the lek paradox. Genetica 132(2):209-216.
  • Heubel KU, Hornhardt K, Ollmann T, Parzefall J, Ryan MJ, Schlupp I (2008): Geographic variation in female mate-copying in the species complex of a unisexual fish, Poecilia formosa. Behaviour 145:1041-1064.
  • Heubel KU, Lindström K, Kokko H (2008): Females increase current reproductive effort when future access to males is uncertain. Biology Letters 4(2):224-227.
  • Kokko H, Heubel KU, Rankin DJ (2008): How populations persist when asexuality requires sex: the spatial dynamics of coping with sperm parasites. Proceedings of the Royal Society B: Biological Sciences 275(1636):817-825.
  • Tobler M, Schlupp I, Heubel KU, Riesch R, de Leon FJG, Giere O, Plath M (2006): Life on the edge: hydrogen sulfide and the fish communities of a Mexican cave and surrounding waters. Extremophiles 10(6):577-585.
  • Plath M, Seggel U, Burmeister H, Heubel KU, Schlupp I (2006) Choosy males from the underground: male mating preferences in surface- and cave-dwelling Atlantic mollies (Poecilia mexicana). Naturwissenschaften 93(3):103-109.
  • Heubel KU, Schlupp I (2006): Turbidity affects association behaviour in male Poecilia latipinna. Journal of Fish Biology 68(2):555-568.
  • Plath M., Heubel K.U., de Leon F.G. & Schlupp I. (2005): Cave molly females (Poecilia mexicana, Poeciliidae, Teleostei) like well-fed males. Behavioral Ecology and Sociobiology, 58(2):144-151.
  • Plath M., Heubel K.U. & Schlupp I. (2005): Field observations on male mating behavior in surface- and cave-dwelling Atlantic mollies (Poecilia mexicana, Poeciliidae). Zeitschrift für Fischkunde, 7:113-119.  Book chapter.
  • Heubel K, Hoffmeister T-S (1999): Emancipated female flies? Mate choice in a lekking fruit fly. Proceedings of the German Society for General and Applied Entomology 12:455-458.